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LoRa/FSK DIO Lines I/O or Out ?

Hi There
I am looking at code which is derived from Semtech, 2013.

In it DIO 0 - 3 are used but treated as input, pull-down by the STM32L0+ processor. Which seems fair enough.

The data sheet describes DIO0-5 as I/O but all the uses are as outputs, push pull.

Do these lines ever change to Input (from the SX1276 point of view) ?

Or are they ever anything other than outputs to the CPU?

Also what happens to the lines during shutdown and standby modes?

Regards
Simon